Professor Citachka · Department
Neuroscience
A clear route, built for real understanding
Your guided study path
Move through sections → levels → units → lessons. Each lesson is deliberately longer than a drill: it explains the idea, works through an example, gives you optional retrieval practice, and then saves your reading. Unit quizzes and level assessments come after the teaching—not every two minutes, because we are building knowledge, not training a very anxious button-pusher.
Your progress path
Units, one crown at a time
Each unit is five lessons. Read every lesson in a unit and it earns a crown. The next unit opens when you reach it, so the path always shows exactly where you are.
- —day streak
- —of 14 unit crowns
- —of 70 lessons read
- —today’s goal: one lesson
You are hereUnit 1: Cells, electrical gradients and synapsesContinue this unit
Section 1Cells to behavior
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Unit 2: Circuit balance, systems and behavior0 of 5 lessons read
Finish Unit 1: Cells, electrical gradients and synapses to open this unit.
Section 2Learning and interpreting evidence
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Unit 3: Plasticity, memory and learning0 of 5 lessons read
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Unit 4: Context and responsible scientific inference0 of 5 lessons read
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Section 3Mechanisms, representations, and the logic of neural evidence
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Unit 5: From synaptic integration to experience-dependent circuits0 of 5 lessons read
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Unit 6: From memory models to defensible neuroscience claims0 of 5 lessons read
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Section 4From transmission to regulation: synapses, systems, and the limits of measurement
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Unit 7: Release, plasticity, and memory across synapses and systems0 of 5 lessons read
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Unit 8: Sensation, action, regulation, and the measurement of brain activity0 of 5 lessons read
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Section 5Cortex, wiring and modulation: from maps to habits and the appraisal of brain claims
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Unit 9: Cortical organisation, development and neuromodulation0 of 5 lessons read
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Unit 10: Networks, habits and the appraisal of evidence0 of 5 lessons read
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Section 6Sensation, regulation and the brain-body boundary: what the pathways do and do not prove
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Unit 11: Sensation as construction: from receptor sheets to perceptual signals0 of 5 lessons read
Finish Unit 1: Cells, electrical gradients and synapses to open this unit.
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Unit 12: Inside the body and outside the moment: regulation, timing, and a contested metaphor0 of 5 lessons read
Finish Unit 1: Cells, electrical gradients and synapses to open this unit.
Section 7Change under scrutiny: how learning, development, ageing and interventions change the brain, and how such claims are tested
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Unit 13: Plasticity in practice: skills, training, development and ageing0 of 5 lessons read
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Unit 14: Mechanisms and interventions put to the test0 of 5 lessons read
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Continue Neuroscience: Neurons and gliaLevel 1 · Topic 1 of 10 · Your next unfinished lesson
Expanded instructional sections: 50 / 70 published course lessons. Original explanations elsewhere remain available but are not counted as expanded. Earlier assessment banks reuse lesson questions; appended course levels have separately authored application questions and longer cumulative tests. Extended lessons and book guides retain separate reading sequences; their own multi-unit assessment maps are not yet authored.
Published now: 14 of 100 requested core units. These are grouped into 7 published core levels, not the full requested curriculum or a professional qualification. The remaining units are not published, not hidden behind a paywall or presented as completed work.
Section 1 · FoundationCells to behavior0/10 lessons read · 2 units
Level 1 · Unit 1
Cells, electrical gradients and synapses
Unit guidebook
Before you begin, this unit teaches Neurons and glia, Membrane gradients, Resting potential, Action potentials, Synaptic transmission. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Neurons and gliaStart lesson →Lesson 2
Membrane gradientsSave the previous lesson to continueLesson 3
Resting potentialSave the previous lesson to continueLesson 4
Action potentialsSave the previous lesson to continueLesson 5
Synaptic transmissionSave the previous lesson to continue
Level 1 · Unit 2
Circuit balance, systems and behavior
Unit guidebook
Before you begin, this unit teaches Excitation and inhibition, Myelin and conduction, Central and peripheral systems, Sensory coding, Motor control. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Excitation and inhibitionSave the previous lesson to continueLesson 2
Myelin and conductionSave the previous lesson to continueLesson 3
Central and peripheral systemsSave the previous lesson to continueLesson 4
Sensory codingSave the previous lesson to continueLesson 5
Motor controlSave the previous lesson to continue
Level 1 assessment
10 questions · Not assessed
Section 2 · DevelopmentLearning and interpreting evidence0/10 lessons read · 2 units
Level 2 · Unit 1
Plasticity, memory and learning
Unit guidebook
Before you begin, this unit teaches Plasticity and timescale, Memory systems, Retrieval and spacing, Attention and limited resources, Reward and prediction error. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Plasticity and timescaleSave the previous lesson to continueLesson 2
Memory systemsSave the previous lesson to continueLesson 3
Retrieval and spacingSave the previous lesson to continueLesson 4
Attention and limited resourcesSave the previous lesson to continueLesson 5
Reward and prediction errorSave the previous lesson to continue
Level 2 · Unit 2
Context and responsible scientific inference
Unit guidebook
Before you begin, this unit teaches Stress and context, Reading a brain image, Correlation and intervention, Reverse inference, Evaluate a neuroscience headline. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Stress and contextSave the previous lesson to continueLesson 2
Reading a brain imageSave the previous lesson to continueLesson 3
Correlation and interventionSave the previous lesson to continueLesson 4
Reverse inferenceSave the previous lesson to continueLesson 5
Evaluate a neuroscience headlineSave the previous lesson to continue
Level 2 assessment
10 questions · Not assessed
Section 3 · IntegrationMechanisms, representations, and the logic of neural evidence0/10 lessons read · 2 units
Level 3 · Unit 1
From synaptic integration to experience-dependent circuits
Unit guidebook
Before you begin, this unit teaches Synaptic integration is a spatiotemporal computation, Inhibition changes responsiveness, not only voltage, Center–surround fields encode contrast, Population codes and ambiguous single-cell responses, NMDA-dependent potentiation: induction versus expression. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Synaptic integration is a spatiotemporal computationSave the previous lesson to continueLesson 2
Inhibition changes responsiveness, not only voltageSave the previous lesson to continueLesson 3
Center–surround fields encode contrastSave the previous lesson to continueLesson 4
Population codes and ambiguous single-cell responsesSave the previous lesson to continueLesson 5
NMDA-dependent potentiation: induction versus expressionSave the previous lesson to continue
Level 3 · Unit 2
From memory models to defensible neuroscience claims
Unit guidebook
Before you begin, this unit teaches Hippocampal indexing: a model across explanatory levels, BOLD, local field potentials, and what a signal measures, Necessity, sufficiency, and competing causal pathways, Reverse inference requires selectivity and a comparison set, Build a multi-level evidence argument without overclaiming. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Hippocampal indexing: a model across explanatory levelsSave the previous lesson to continueLesson 2
BOLD, local field potentials, and what a signal measuresSave the previous lesson to continueLesson 3
Necessity, sufficiency, and competing causal pathwaysSave the previous lesson to continueLesson 4
Reverse inference requires selectivity and a comparison setSave the previous lesson to continueLesson 5
Build a multi-level evidence argument without overclaimingSave the previous lesson to continue
Level 3 assessment
12 questions · Not assessed
Section 4 · IntegrationFrom transmission to regulation: synapses, systems, and the limits of measurement0/10 lessons read · 2 units
Level 4 · Unit 1
Release, plasticity, and memory across synapses and systems
Unit guidebook
Before you begin, this unit teaches Release probability, quantal size, and how a synapse reports uncertainty, The postsynaptic side: receptor complement, driving force, and excitatory-inhibitory balance, Plasticity runs both ways: depression, synaptic scaling, and metaplasticity, Consolidation, reactivation, and reconsolidation: the standard model and its disputes, Sleep, oscillations, and memory: the active consolidation hypothesis and its limits. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Release probability, quantal size, and how a synapse reports uncertaintySave the previous lesson to continueLesson 2
The postsynaptic side: receptor complement, driving force, and excitatory-inhibitory balanceSave the previous lesson to continueLesson 3
Plasticity runs both ways: depression, synaptic scaling, and metaplasticitySave the previous lesson to continueLesson 4
Consolidation, reactivation, and reconsolidation: the standard model and its disputesSave the previous lesson to continueLesson 5
Sleep, oscillations, and memory: the active consolidation hypothesis and its limitsSave the previous lesson to continue
Level 4 · Unit 2
Sensation, action, regulation, and the measurement of brain activity
Unit guidebook
Before you begin, this unit teaches Transduction and adaptation: turning physical energy into spike trains, Motor units, loops, and error correction: how the nervous system produces movement, Autonomic control and homeostasis: reflex loops, feedback, and the limits of a set point, Electroencephalography and magnetoencephalography: millisecond timing with blurred location, Functional imaging, lesions, and causal inference: choosing a method that can bear the claim. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Transduction and adaptation: turning physical energy into spike trainsSave the previous lesson to continueLesson 2
Motor units, loops, and error correction: how the nervous system produces movementSave the previous lesson to continueLesson 3
Autonomic control and homeostasis: reflex loops, feedback, and the limits of a set pointSave the previous lesson to continueLesson 4
Electroencephalography and magnetoencephalography: millisecond timing with blurred locationSave the previous lesson to continueLesson 5
Functional imaging, lesions, and causal inference: choosing a method that can bear the claimSave the previous lesson to continue
Level 4 assessment
12 questions · Not assessed
Section 5 · IntegrationCortex, wiring and modulation: from maps to habits and the appraisal of brain claims0/10 lessons read · 2 units
Level 5 · Unit 1
Cortical organisation, development and neuromodulation
Unit guidebook
Before you begin, this unit teaches Cortical maps and topographic organisation: what a map claim requires, How experience wires circuits: pruning, critical periods, and two kinds of plasticity, Dopamine: prediction error, effort, and why 'the pleasure molecule' is a misreading, Stress physiology and its cumulative effects on brain and body, Sleep as a controlled system: two processes, arousal circuits, and what sleep changes. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Cortical maps and topographic organisation: what a map claim requiresSave the previous lesson to continueLesson 2
How experience wires circuits: pruning, critical periods, and two kinds of plasticitySave the previous lesson to continueLesson 3
Dopamine: prediction error, effort, and why 'the pleasure molecule' is a misreadingSave the previous lesson to continueLesson 4
Stress physiology and its cumulative effects on brain and bodySave the previous lesson to continueLesson 5
Sleep as a controlled system: two processes, arousal circuits, and what sleep changesSave the previous lesson to continue
Level 5 · Unit 2
Networks, habits and the appraisal of evidence
Unit guidebook
Before you begin, this unit teaches Language and its lateralisation: dissociations, networks, and the limits of left-right claims, Executive function and the limits of grey-matter claims, The biology of habit: goal-directed and habitual control, Reading a neuroscience headline: correlation, causation, and replication, Capstone: appraising a causal claim about the brain, from design to headline. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Language and its lateralisation: dissociations, networks, and the limits of left-right claimsSave the previous lesson to continueLesson 2
Executive function and the limits of grey-matter claimsSave the previous lesson to continueLesson 3
The biology of habit: goal-directed and habitual controlSave the previous lesson to continueLesson 4
Reading a neuroscience headline: correlation, causation, and replicationSave the previous lesson to continueLesson 5
Capstone: appraising a causal claim about the brain, from design to headlineSave the previous lesson to continue
Level 5 assessment
Section 6 · IntegrationSensation, regulation and the brain-body boundary: what the pathways do and do not prove0/10 lessons read · 2 units
Level 6 · Unit 1
Sensation as construction: from receptor sheets to perceptual signals
Unit guidebook
Before you begin, this unit teaches The visual system as a worked example of hierarchical processing, Hearing: how the auditory system extracts pitch and location, Touch: mechanoreceptors, submodalities and two rival coding schemes, Pain is a constructed signal, not a direct readout of tissue damage, Vestibular and proprioceptive sense of self-position. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
The visual system as a worked example of hierarchical processingSave the previous lesson to continueLesson 2
Hearing: how the auditory system extracts pitch and locationSave the previous lesson to continueLesson 3
Touch: mechanoreceptors, submodalities and two rival coding schemesSave the previous lesson to continueLesson 4
Pain is a constructed signal, not a direct readout of tissue damageSave the previous lesson to continueLesson 5
Vestibular and proprioceptive sense of self-positionSave the previous lesson to continue
Level 6 · Unit 2
Inside the body and outside the moment: regulation, timing, and a contested metaphor
Unit guidebook
Before you begin, this unit teaches Movement is planned before it is executed, Interoception and the immune-to-brain conversation, How the brain regulates temperature, hunger and thirst, Circadian biology beyond sleep: an internal clock that times many systems, Capstone: is the brain a computer, and where does the metaphor break?. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Movement is planned before it is executedSave the previous lesson to continueLesson 2
Interoception and the immune-to-brain conversationSave the previous lesson to continueLesson 3
How the brain regulates temperature, hunger and thirstSave the previous lesson to continueLesson 4
Circadian biology beyond sleep: an internal clock that times many systemsSave the previous lesson to continueLesson 5
Capstone: is the brain a computer, and where does the metaphor break?Save the previous lesson to continue
Level 6 assessment
Section 7 · IntegrationChange under scrutiny: how learning, development, ageing and interventions change the brain, and how such claims are tested0/10 lessons read · 2 units
Level 7 · Unit 1
Plasticity in practice: skills, training, development and ageing
Unit guidebook
Before you begin, this unit teaches What a skill is, neurally: components, stages and why 'the skill area' is a category error, Training-induced change and everyday plasticity claims are different kinds of evidence, Critical periods revisited: what vision and language evidence actually constrains, Adolescent brain development: what the imaging shows, and what the 'immature brake' story adds, Ageing: separating normal cognitive change from disease, and why the boundary is a measurement decision. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
What a skill is, neurally: components, stages and why 'the skill area' is a category errorSave the previous lesson to continueLesson 2
Training-induced change and everyday plasticity claims are different kinds of evidenceSave the previous lesson to continueLesson 3
Critical periods revisited: what vision and language evidence actually constrainsSave the previous lesson to continueLesson 4
Adolescent brain development: what the imaging shows, and what the 'immature brake' story addsSave the previous lesson to continueLesson 5
Ageing: separating normal cognitive change from disease, and why the boundary is a measurement decisionSave the previous lesson to continue
Level 7 · Unit 2
Mechanisms and interventions put to the test
Unit guidebook
Before you begin, this unit teaches Memory reconsolidation: the biology, the boundary conditions, and why the debate is unresolved, Attention as selection, and what the evidence about media multitasking actually shows, Habit and intention: two control systems, how they interact, and what 'habit' does not mean, Pharmacological and behavioural interventions: what each kind of trial can and cannot show, Capstone: evaluating a commercial brain-training claim end to end. The lessons introduce ideas in order, then the unit quiz asks you to retrieve and apply them. You may revisit completed lessons whenever you wish.
Lesson 1
Memory reconsolidation: the biology, the boundary conditions, and why the debate is unresolvedSave the previous lesson to continueLesson 2
Attention as selection, and what the evidence about media multitasking actually showsSave the previous lesson to continueLesson 3
Habit and intention: two control systems, how they interact, and what 'habit' does not meanSave the previous lesson to continueLesson 4
Pharmacological and behavioural interventions: what each kind of trial can and cannot showSave the previous lesson to continueLesson 5
Capstone: evaluating a commercial brain-training claim end to endSave the previous lesson to continue
Level 7 assessment
Progress stays in this browser, on this device—not an account or cloud backup. Clearing browser data removes it. Visits never count as study. Previous untimed legacy completions are not invented as study days.
From neurons and synapses to cognition, emotion, and behavior.
The nervous system rewards patient study: structure first, mechanism second, interpretation last.
Your pathway
Study Map
Each unit builds on the one before it. We advance through demonstrated understanding, not mere completion.
- 01
Neural foundations
Cells, electrical signaling, synapses, and the organization of the nervous system.
Begin Unit 1: Neural Foundations - 02
Sensation and movement
How brains receive information, construct perception, and guide action.
Continue to Lesson 02: Sensation and movement - 03
Learning, memory, and attention
Plasticity, memory systems, attention, and the science of practice.
Continue to Lesson 03: Learning, memory, and attention - 04
Emotion and motivation
Stress, reward, affective circuits, and decision-making.
Continue to Lesson 04: Emotion and motivation - 05
Clinical and cognitive neuroscience
Brain injury, neurodevelopment, psychiatric frameworks, and responsible interpretation.
Continue to Lesson 05: Clinical and cognitive neuroscience
Beyond the foundation
Your expanding curriculum
This department does not stop at beginner level. Each stage adds complexity, independence, and real-world application.
- Level 1Foundation: cells, circuits, and core brain systems
- Level 2Core: perception, movement, memory, and emotion
- Level 3Intermediate: systems neuroscience and experimental reasoning
- Level 4Advanced: clinical neuroscience, papers, and research methods
- Level 5Independent mastery: critically synthesize new evidence